Adam Tofilski

Adam Tofilski

professor at University of Agriculture in Krakow

Publications

  1. Bakhchou S., Aglagane A., Tofilski A., Mokrini F., Er-Rguibi O., El Mouden E.H., Machlowska J., Fellahi S., Mohssine E.H. (2025). Morphological diversity of Moroccan honey bees (Apis mellifera L. 1758): insights from a geometric morphometric study of wing venation in honey bees from different climatic regions. Diversity 17(8):527. HLML

  2. Payró de la Cruz E., Valencia Domínguez M., Ramos Reyes R., Tofilski A. (2025). Reexamination of honey bee Africanization in Mexico and other regions of the New World. Scientific Reports 15(1): 1-10. HLML

  3. Ganie S.A., Rehman S.A., Bano P., Tofilski A. (2025). Influence of host plant on the wing shape of Bactrocera cucurbitae and Bactrocera tau (Diptera: Tephritidae). Oriental Insects 59:19-32. HLML

  4. Kaur H., Ganie S.A., Tofilski A. 2024. Morphometric identification of an unknown honey bee colony: an example from north India. Journal of Apicultural Science 68:155-166. https://doi.org/10.2478/JAS-2024-0013

  5. Kaur H., Nedić N., Tofilski A. 2024. Influence of honey bee (Apis mellifera) breeding on wing venation in Serbia and neighbouring countries. PeerJ 12:e17247 HLML

  6. Tofilski A., Kaur H., Łopuch S. 2024. Size and shape differences in the fore wings of honey bee (Apis mellifera) queens, workers, and drones. Journal of Apicultural Science 68:5-17. HLML

  7. Temirbayeva, K., Torekhanov A., Nuralieva U., Sheralieva Z., Tofilski A. 2023. In Search of Apis mellifera pomonella in Kazakhstan. Life 13:1860. HLML

  8. Aglagane A., Oleksa A., Er-Rguibi O., Tofilski A., Laghzaoui E.-M., El Mouden E.H., Aamiri A., Aourir M. 2023. Genetic diversity and population structure of the Saharan honey bee Apis mellifera sahariensis from southeastern Morocco: introgression assessment and implications for conservation. Apidologie 54:31 HLML

  9. Oleksa, A., Căuia, E., Siceanu, A., Puškadija, Z., Kovačić, M., Pinto, M. A., Rodrigues, P. J., Hatjina, F., Charistos, L., Bouga, M., Prešern, J., Kandemir, İ., Rašić, S., Kusza, S., Tofilski, A. 2023. Honey bee (Apis mellifera) wing images: a tool for identification and conservation. GigaScience, 12: giad019. HLML

  10. García C.A.Y., Rodrigues P.J., Tofilski A., Elen D., McCormak G.P., Oleksa A., Henriques D., Ilyasov R., Kartashev A., Bargain C., Fried B., Pinto M.A. 2022. Using the Software DeepWings© to Classify Honey Bees across Europe through Wing Geometric Morphometrics. Insects, 13: 1132. HLML

  11. Szpila K., Johnston N. P., Akbarzadeh K., Richet R., Tofilski A. 2022. Wing measurements are a possible tool for the identification of European forensically important Sarcophagidae. Forensic Science International, 340: 111451. HLML

  12. Aglagane A., Tofilski A., Er-Rguibi O., Laghzaoui E.-M., Kimdil L., El Mouden E.H., Fuchs S., Oleksa A., Aamiri A., Aourir M. 2022. Geographical variation of honey bee (Apis mellifera L. 1758) populations in South-Eastern Morocco: a geometric morphometric analysis. Insects 13, 288. HLML

  13. Dukku U. H., Fuchs S., Danailu G., Grünewald B., Tofilski A., Kryger P., Meixner M. D. 2022. Morphometric and mitochondrial variation of Apis mellifera L. and its relationship with geographical variables in parts of West and Central Africa. Journal of Apicultural Research, HLML

  14. Łopuch S., Tofilski A. 2021. The effect of a lack of uncapped brood on social interactions between honey bee workers and the queen. Apidologie, 52:1346-1357. HLML VIDEO

  15. Janczyk A., Tofilski A., 2021. Monthly changes in honey bee forewings estimated using geometric morphometrics, Journal of Apicultural Science 65:139 - 146. HLML

  16. Tofilski A., Căuia E., Siceanu A., Vișan G.O., Căuia D. 2021. Historical changes in honey bee wing venation in Romania. Insects 12: 542. HLML

  17. Oleksa A., Kusza S. Tofilski A. 2021. Mitochondrial DNA Suggests the introduction of honeybees of African ancestry to East-Central Europe. Insects 12: 410. HLML

  18. Puškadija Z., Kovačić M., Raguž N., Lukić B., Prešern J., Tofilski A. 2021. Morphological diversity of Carniolan honey bee (Apis mellifera carnica) in Croatia. Journal of Apicultural Research 60: 326-336. HLML

  19. Janczyk A., Meixner M.D., Tofilski A. 2021. Morphometric identification of the endemic Maltese honey bee (Apis mellifera ruttneri). Journal of Apicultural Research 60: 157-164. HLML

  20. Hailu T. G., D’Alvise P., Tofilski A., Fuchs S., Greiling J., Rosenkranz P., Hasselmann M. (2020). Insights into Ethiopian honey bee diversity based on wing geomorphometric and mitochondrial DNA analyses. Apidologie 51: 1182-1198. HLML

  21. Goczał J., Oleksa A., Rossa R., Chybicki I., Meyza K., Plewa R., Landvik M., Gobbi M., Hoch G., Tamutis V., Balalaikins M., Telnov D., Dascalu M., Tofilski A. 2020. Climatic oscillations drive plants-insects co-evolution: Divergence patterns across the distribution range of Monochamus sartor reflect Quaternary history of its host tree. Scientific Reports 10: 16524. HLML

  22. Czekońska K., Tofilski A. 2020. Body mass of honey bee drones developing in constant and in changing temperatures. Apidologie, 51: 510-518. HLML

  23. Łopuch S., Tofilski A. 2020. Impact of the quality of food sources on the wing beating of honey bee dancers. Apidologie 51: 631-641. HLML

  24. Czekońska K., Szentgyörgyi H., Tofilski A. 2019. Body mass but not wing size or symmetry correlates with life span of honey bee drones. Bulletin of Entomological Research 109: 383-389. HLML

  25. Łopuch S., Tofilski A. 2019. Use of high-speed video recording to detect wing beating produced by honey bees. Insectes Sociaux, 66: 235-244. HLML

  26. Szpila K., Żmuda A., Akbarzadeh K., Tofilski A. 2019. Wing measurement can be used to identify European blow flies (Diptera: Calliphoridae) of forensic importance. Forensic Science International, 296: 1-8. HLML

  27. Goczał J., Rossa R., Tofilski A., 2019. Intersexual and intrasexual patterns of horn size and shape variation in the European rhinoceros beetle: quantifying the shape of weapons. Biological Journal of the Linnean Society, 127: 34-43. HLML

  28. Węgrzynowicz P., Gerula D., Tofilski A., Panasiuk B., Bieńkowska M., 2019. Maternal inheritance in hybrids of three honey bee subspecies. Journal of Apicultural Science, 63: 131-138. HLML

  29. Szentgyörgyi H., Czekońska K., Tofilski A. 2018 Honey bees are larger and live longer after developing at low temperature. Journal of Thermal Biology 78: 219-226. HLML

  30. Tofilski A., Chuda-Mickiewicz B., Czekońska K. 2018. Circular movement of honey bee spermatozoa inside spermatheca. Invertebrate Reproduction & Development 62: 63-66. HLML

  31. Goczał J., Rossa R., Nawrocka A., Sweeney J.D., Tofilski A. 2018. Developmental costs of biological invasion: The exotic wood borer Tetropium fuscum (Coleoptera: Cerambycidae) is more asymmetric and smaller in invaded area. Environmental Entomology 47: 982-989. HLML

  32. Mielczarek Ł.E., Tofilski A. 2018. Semiautomated identification of a large number of hoverfly (Diptera: Syrphidae) species based on wing measurements. Oriental Insects 52: 245-258. HLML

  33. Nawrocka A., Kandemir I., Fuchs S., Tofilski A. 2018. Computer software for identification of honey bee subspecies and evolutionary lineages. Apidologie 49: 172-184. HLML

  34. Goczał J., Rossa R., Tofilski A. 2018. Elytra reduction may affect the evolution of beetle hind wings. Zoomorphology 137: 131-138. HLML

  35. Tofilski A. 2018. DKey software for editing and browsing dichotomous keys. ZooKeys 735: 131-140. HLML

  36. Łopuch S., Tofilski A. 2017. Importance of wing movements for information transfer during honey bee waggle dance. Ethology 123: 974-980. HLML

  37. Szentgyorgyi H., Moroń D., Nawrocka A., Tofilski A., Woyciechowski M. 2017. Forewing structure of the solitary bee Osmia bicornis developing on heavy metal pollution gradient. Ecotoxicology 26: 1031-1040. HLML

  38. Szentgyörgyi H., Czekońska K., Tofilski A. 2017. The effects of starvation of honey bee larvae on reproductive quality and wing asymmetry of honey bee drones. Journal of Apicultural Science 61: 233-243. HLML

  39. Goczał J., Rossa R., Sweeney J., Tofilski A. 2017. Citizen monitoring of invasive species: wing morphometry as a tool for detection of alien Tetropium species. Journal of Applied Entomology 141: 496-506. HLML

  40. Grzywacz A., Ogiela J., Tofilski A. 2017. Identification of Muscidae (Diptera) of medico-legal importance by means of wing measurements. Parasitology Research 116: 1495-1504. HLML

  41. Łopuch S., Tofilski A. 2017. Direct visual observation of wing movements during the honey bee waggle dance. Journal of Insect Behavior 30: 199-210. HLML

  42. Rossa R., Goczał J., Tofilski A. 2017. Hind wing morphology facilitates discrimination between two sibling species: Leiopus nebulosus and L. linnei (Coleoptera: Cerambycidae). Zootaxa 4227: 266-278. HLML

  43. Szentgyörgyi H., Czekońska K., Tofilski A. 2016. Influence of pollen deprivation on the fore wing asymmetry of honeybee workers and drones. Apidologie 47: 653-662. HLML

  44. Łopuch S., Tofilski A. 2016. The relationship between asymmetry, size and unusual venation in honey bees (Apis mellifera). Bulletin of Entomological Research 106: 304-313. HLML

  45. Przybyłowicz Ł., Pniak M., Tofilski A. 2016. Semiautomated Identification of European Corn Borer (Lepidoptera: Crambidae). Journal of Economic Entomology 109:195-199. HLML

  46. Rossa R., Goczał J., Tofilski A. 2016. Within- and between-species variation of wing venation in genus Monochamus (Coleoptera: Cerambycidae). Journal of Insect Science 16:5. HLML

  47. Mielczarek L.E., Oleksa A., Meyza K., Tofilski A. 2016. Seasonal polyphenism in Eristalis pertinax (Diptera: Syrphidae). European Journal of Entomology 113:489-496. HLML

  48. Oleksa A., Tofilski A. 2015. Wing geometric morphometrics and microsatellite analysis provide similar discrimination of honey bee subspecies. Apidologie 46: 49-60. HLML

  49. Tofilski A. 2014. A scientific note on amoeboid movement of honey bee semen. Apidologie 45:637-640. HLML

  50. Soszyński B., Mielczarek L.E., Tofilski A. 2013. Dasysyrphus neovenustus sp. n. (Diptera: Syrphidae) a new species in the venustus species group. Polish Journal of Entomology 82:353-363. HLML

  51. Wenseleers T., Bacon J.P., Alves D.A., Couvillon M.J., Kärcher M., Nascimento F.S., Nogueira-Neto P., Ribeiro M., Robinson E.J.H., Tofilski A., Ratnieks F.L.W. 2013. Bourgeois behavior and freeloading in the colonial orb web spider Parawixia bistriata (Araneae, Araneidae). American Naturalist 182:120-129. HLML

  52. Oleksa A., Wilde J., Tofilski A., Chybicki I.J. 2013. Partial reproductive isolation between European subspecies of honey bees. Apidologie 44:611-619. HLML

  53. Tofilski A., Oleksa A. 2013. There are still bee trees in Europe. Bee World 90:18-19. HLML

  54. Oleksa A., Gawroński R., Tofilski A. 2013. Rural avenues as a refuge for feral honey bee population. Journal of Insect Conservation 17:465-472. HLML

  55. Tofilski A., Chuda-Mickiewicz B., Czekonska K., Chorbiński P. 2012. Flow cytometry evidence about sperm competition in honey bee (Apis mellifera). Apidologie 43: 63-70. HLML

  56. Tofilski A. 2011. Homology of submarginal cross veins in forewings of bees (Hymenoptera: Apiformes). Journal of Apicultural Science 55:131-140. PDF

  57. Higginson A. D., Barnard C.J., Tofilski A., Medina L., Ratnieks F.L.W., 2011. Experimental wing damage affects foraging effort and foraging distance in honeybees Apis mellifera. Psyche 2011:419793. HLML

  58. Oleksa A., Chybicki I., Tofilski A., Burczyk J., 2011. Nuclear and mitochondrial patterns of introgression into native dark bees (Apis mellifera mellifera) in Poland. Journal of Apicultural Research 50:116-129. HLML

  59. Bonckaert W., Tofilski A., Nascimento F., Billen J., Ratnieks F., Wenseleers T., 2011. Co-occurrence of three types of egg policing in the Norwegian wasp Dolichovespula norwegica. Behavioral Ecology and Sociobiology 65:633-640. HLML

  60. Tofilski A., 2009. Shorter-lived workers start foraging earlier. Insectes Sociaux 56:359-366. HLML

  61. Gerula D., Tofilski A., Węgrzynowicz P., Skowronek W., 2009. Computer-assisted discrimination of honeybee subspecies used for breeding in Poland. Journal of Apicultural Science 53:105-114.

  62. Tofilski A., 2008. Using geometric morphometrics and standard morphometry to discriminate three honeybee subspecies. Apidologie 39:558-563. HLML

  63. Tofilski A., Couvillon M.J., Evison S.E.F., Helanterä H., Robinson E.J.H., Ratnieks F.L.W., 2008. Preemptive defensive self-sacrifice by ant workers. American Naturalist 172:E239-E243. HLML

  64. Robinson E.J.H., Tofilski A., Ratnieks F.L.W., 2008. The use of native and non-native tree species for foraging and nesting habitat by the wood-ant Formica lugubris (Hymenoptera: Formicidae). Myrmecological News 11:1-7.

  65. Tofilski A., Ratnieks F.L.W. 2008. Simple rules based on pile slope are used in the self organization of sand pile formation by Pheidole oxyops ants. Insectes Sociaux 55:37-42. HLML

  66. Tofilski A. 2007. Automatic measurements of honeybee wings. In MacLeod, N. (ed.), Automated Object Identification in Systematics: Theory, Approaches, and Applications, Systematics Association Special Volume, CRC Press, Boca Raton, Florida, pp 289-298. PDF

  67. Tofilski A. 2006. Influence of caste polyethism on longevity of workers in social insect colonies. J. Theor. Biol. 238:527-531. PDF, Appendix

  68. D’Ettorre P., Tofilski A., Heinze J., Ratnieks F. L. W., 2006. Non-transferable signals on ant queen eggs. Naturwissenschaften 93:136-140.

  69. Helanterä H., Tofilski A., Wenseleers T., Ratnieks F. L. W., 2006. Worker policing in the common wasp Vespula vulgaris is not aimed at improving colony hygiene. Insectes Sociaux 53:399-402.

  70. Tofilski A., Ratnieks F.L.W. 2005. Sand pile formation in Dorymyrmex ants. Journal of Insect Behavior 18:505-512. PDF

  71. Wenseleers T, Tofilski A, Ratnieks FLW, 2005. Queen and worker policing in the tree wasp Dolichovespula sylvestris. Behavioral Ecology and Sociobiology 58:80-86. PDF

  72. Wenseleers T, Badcock NS, Erven K, Tofilski A, Nascimento FS, Hart AG, Burke TA, Archer ME, Ratnieks FLW, 2005. A test of worker policing theory in an advanced eusocial wasp, Vespula rufa. Evolution 59:1306-1314. PDF

  73. Nascimento F. S., Hrncir M., Tofilski A., Zucchi R., 2005. Scraping sounds produced by a social wasp (Asteloeca ujhelyii, Hymenoptera: Vespidae). Ethology 111:1116-1125.

  74. Jensen A., Palmer K., Chaline N., Raine N., Tofilski A., Martin S., Pedersen B., Boomsma J., Ratnieks F., 2005. Quantifying honey bee mating range and isolation in semi-isolated valleys by DNA microsatellite paternity analysis. Conservation Genetics 6:527-537.

  75. Tofilski A. 2004. DrawWing, a program for numerical description of insect wings. Journal of Insect Science, 4:17, 5pp. HLML

  76. Tofilski A., Czekonska K. 2004. Emergency queen rearing in honeybee colonies with brood of known age. Apidologie 35:275-282. PDF

  77. Tofilski A. 2002. Influence of age polyethism on longevity of workers in social insects. Behav. Ecol. Sociobiol. 51: 234-237. PDF, Appendix

  78. Tofilski A. 2000. Senescence and learning in honey bee (Apis mellifera) workers. Acta. Neurobiol. Exp. 60:35-39. PDF

  79. Tofilski A., Kopel J. 1996. The influence of Nosema apis on maturation and flight activity of honey bee drones. Pszczelnicze Zeszyty Naukowe 40:55-60. PDF

  80. Tofilski A. 1995. When should honey bee (Apis mellifera) colonies swarm? I Krajowa Konferencja Zastosowań Matematyki w Biologii i Medycynie, Zakopane. PDF